1992 - Professor of Molecular Ecology Department of Biology, University of York 1979 - 1992 Research group leader John Innes Institute, Norwich 1978 - 1979 SRC Postdoctoral Fellow School of Biology, University of Sussex 1975 - 1978 NIH Postdoctoral Fellow Department of Ecology and Evolution, State University of New York at Stony Brook 1975 PhD University of Cambridge (Natural Sciences - Genetics) 1972 BA University of Cambridge (Natural Sciences - Genetics)