1984 – 1987, Dept. of Biochemistry, Stanford University. Research topic: Enzymes of DNA recombination in E. coli. Ph. D., Biochemistry, 1984, University of California, Berkeley. Dissertation topic: Enzyme mechanisms and kinetic isotope effects. B.A., Chemistry, 1978, Haverford College, Haverford, PA.

Enzymology and mechanisms of DNA recombination and repair. Mechanism of the RecBCD enzyme from E. coli. DNA repair in Deinococcus radiodurans