BSc (Hons) University of Leeds (1982). PhD University of Manchester (1986). Associate Research Scientist Oklahoma Medical Research Foundation, USA (1987). MRC Postdoctoral Research Fellow University of Cambridge (1987-1990). Lecturer, University of Wales College of Medicine (1990-1994). Lecturer, Senior Lecturer, Reader, University of Leicester (1994-2010). Visiting Scientist, Harvard University & Beth Israel Deaconess Medical Centre, USA (1998). Professor of Cell Signalling University of Leicester (2010- ). Visiting Scientist, MRC Laboratory of Molecular Biology, Cambridge (2011- 2012).

How receptors work: mechanistic, functional, and structural biology of receptors Directed evolution for creating new proteins and understanding protein function and structure