2013-present University of Edinburgh Royal Society University Research Fellowship 2012-2013 University of Oxford Royal Society University Research Fellowship 2008-2011 University of Edinburgh NERC Post-Doctoral Research Fellowship 2007-2008 University of Edinburgh Leverhulme Post-Doc with Loeske Kruuk 2005-2006 University of Sheffield NERC Post-Doc with Terry Burke 2001-2005 Imperial College London NERC PhD with Ian Owens 1993-1996 University of Leeds BSc Zoology