1980-1983 PhD in Immunology; Dept. of Immunology, University of Birmingham 1983-1985 Postdoc; Dept. of Immunology, University of Birmingham 1985-1991 Scientific Member, Basel Institute for Immunology, Basel, Switzerland 1991-1998 Wellcome Trust Senior Research Fellow and Honorary Reader/Professor of Cellular Immunology, Royal Postgraduate Medical School, Hammersmith Hospital, London 1998-2002 Wellcome Trust Senior Research Fellow and University Professorial Fellow, ICAPB, University of Edinburgh 2002-Present Professor of Immunology, IIIR, University of Edinburgh