2011 - Professor of Tissue Stem Cell Biology, MRC Centre for Regenerative Medicine, The University of Edinburgh, UK January 2009 – August 2013 Director of the Graduate School of Biological Sciences 1997 – 2011 WT Career Development Fellow, LLR Senior Fellow, then LLR Senior Lecturer, Centre for Genome Research / Institute for Stem Cell Research / MRC Centre for Regenerative Medicine, The University of Edinburgh, UK 1995-1997 WT Advanced Training Fellow, then WT Career Development Fellow, Nuffield Department of Clinical Medicine, University of Oxford 1991-1994 WT Advanced Training Fellow, Walter and Eliza Hall Institute for Medical Research, Melbourne, Australia (with Grant Morahan and Jacques Miller) PhD Biochemistry Imperial College London 1991