2012-present University of Edinburgh Chancellor's Fellow 2011-2012 University of Dundee Principle Investigator 2004-2010 University of Edinburgh Postdoctoral Research Associate in the lab of Prof. Robin Allshire 2000-2004 The Sainsbury Laboratory PhD "Molecular characterisation of a viral suppressor of RNAi" supervised by Prof. Sir David Baulcombe 1997-2000 University of Cambridge B.A. (Hons) Natural Sciences (Part II Plant Sciences)