Brakefield, P.M., Gates, J., Keys, D., Kesbeke, F., Wijngaarden, P.J., Monteiro, A., French, V., and Carroll, S.B. (1996). Development, plasticity and evolution of butterfly eyespot patterns. Nature 384, 236-242. Beldade, P., Koops, K., and Brakefield, P.M. (2002). Developmental constraints versus flexibility in morphological evolution. Nature 416, 844-847. Brakefield, P.M., and Reitsma, N. (1991). Phenotypic Plasticity, Seasonal Climate and the Population Biology of Bicyclus Butterflies (Satyridae) in Malawi. Ecol. Entomol. 16, 291-303. Saccheri, I.J., Brakefield, P.M., and Nichols, R.A. (1996). Severe inbreeding depression and rapid fitness rebound in the butterfly Bicyclus anynana (Satyridae). Evolution 50, 2000-2013. Joron, M., and Brakefield, P.M. (2003). Captivity masks inbreeding effects on male mating success in butterflies. Nature 424, 191-194.