1996-2007 - NERC Centre for Ecology and Hydrology, Banchory 1995-1996 - The Macaulay Land Use Research institute, Aberdeen 1991-1995 - University of Oxford, based at The Institute of Terrestrial Ecology,Banchory

Functional connectivity in fragmented landscapes Range shifting in response to environmental change Modelling dispersal and its demographic consequences Development and application of individual-based simulation models