1996 PhD Microbiology, University of Cape Town 1996-1998 Post-Doctoral Research Fellow, University of Stellenbosch 1999-2003 Wellcome Trust Travelling Fellow, University of Oxford. 2004-2009 Wellcome Trust International Senior Research Fellow, University of Cape 2009- 6th Century Chair in Immunology, University of Aberdeen 2009- Honorary Professor, University of Cape Town 2010- Fellow of the Society of Biology 2011- Fellow of the American Academy of Microbiology 2012- Fellow of the Royal Society of South Africa 2013- Fellow of the Royal Society of Edinburgh 2016- Fellow of the Academy of Medical Sciences