BSc Honours (Botany; Plant Physiology), University of Liverpool 1975-1978

PhD in Environmental Plant Physiology/Plant Ecology, University of Sheffield (NERC unit of Comparative Plant Ecology and ARC Letcombe Laboratory) 1978-1981

Post-Doctoral Research Officer, Birkbeck College, University of London 1981-1983

Lecturer/Senior Lecturer in Plant Biology, Portsmouth Polytechnic/University of Portsmouth 1983-

Chartered Biologist ~1983-

Plant (physiological) ecology; water relations and mineral nutrition, photosynthesis, assimilate partitioning and growth. Root/shoot, source/sink relationships.

Quantitative analysis of plant growth and activity.