B.A., 1981, Williams College M.S., 1985, Yale School of Forestry & Environmental Studies Ph.D., 1990, Yale University

Factors driving "bark-stripping" in Rocky Mountain bristlecone pinecanopy uptake and allocation of nitrogen by conifers, ecophysiology of red cedar, effect of the invasive Amur honeysuckle on forest understory herbs, effects of calcium deficiency on tree ecophysiology, development of Fuzzy Set Ordination (FSO) for use with plant community data.