B.Sc. (Honours) Biology, Queen's University, Kingston, Canada M.A. Biology, University of California, Riverside Ph.D. Biology, University of California, Riverside

Costs and benefits of sexually selected displays; Aggressive intermale interactions; Costs of mating and resulting trade-offs; Evolution in poeciliid fishes, especially the Cuban species Girardinus metallicus; Phenotypic plasticity and geographic variation in mating tactics; Host-parasite interactions.