B.S. in Environmental Science, Johnson State College, 1978 M.S. Plant in Pathology, University of Massachusetts, 1981 Ph.D. in Botany, Washington State University, 1990 (under R.N. Mack), Dissertation Title: Multiple Introduction and Founder Effects in Bromustectorum. An Analysis of Eurasian and North American Populations Postdoctoral Research Associate, Department of Botany, Washington State University (under P.S. Soltis) 1990-1992

Plant population ecology and evolutionary biology; investigations of the factors that influence the level and structure of genetic variation: introduction events, founder effects, gene flow, and mating systems; polyploid speciation and ecological consequences of polyploidy.